Fortin, S. G., Sun X., Jayakumar, A., Ward, B. B, Nitrite-oxidizing bacteria adapted to low-oxygen conditions dominate nitrite oxidation in marine oxygen minimum zones, The ISME Journal, 18:1 https://doi.org/10.1093/ismejo/wrae160 (2024)

Frey, C., Tang, W., Ward, B. B., and Lehmann, M. Sample preservation methods for nitrous oxide concentration and isotope ratio measurements in aquatic environments. Limnology and Oceanography Methods. DOI:10.1002/lom3.10638 (2024)

Tang, W., Talbott, J., Jones, T., Ward, B. B. Effluents of wastewater treatment plants are overlooked sources of N₂O to the atmosphere. Biogeosciences. DOI:10.5194/egusphere-2024-638 (2024)

Tang, W., Fortin, S. G., Intrator, N., Lee, J. A., Kunes, M. A., Jayakumar, A., and Ward, B.B. Determination of site-specific nitrogen cycle reaction kinetics allows accurate simulation of in situ nitrogen transformation rates in a large North American estuary. Limnology and Oceanography. DOI:10.1002/lno.12628 (2024)

Zhao, R., Zhang, I. H., Jayakumar, A., Ward, B. B. and Babbin, A. R. Origin, age, and metabolisms of dominant anammox bacteria in the global oxygen deficient zones. ISME Communications. DOI:10.1093/ismeco/ycae060 (2024)

Intrator, N., Jayakumar, A., and Ward, B.B. Nitrous Oxide Reductase gene (nosZ) Phylogeny and Aquatic Biogeography. Frontiers in Microbiology. 15. DOI:10.3389/fmicb.2024.1407573 (2024)

Qin, W., Wei, S., Zheng, Y., Choi E., Li. X. , Juliet Johnston, J., Wan, X., Abrahamson, B., Flinkstrom, Z., Wang, B., Li, H., Hou, L., Sun, X., Wells, M., Ngo, L., Hunt, K., Urakawa, H., Tao, X., Wang, D., Wang, D., Pan, C., Weber, P., Jiang, J., Zhou, J., Zhang, Y., Stahl, D., Ward, B. B., Mayali, X., Martens-Habbena, W., Winkler, M.-K. Differential substrate affinity and catabolite repression enable preferential use of urea by ammonia-oxidizing bacteria. Nature Microbiology. 9(2), 524-536. DOI:10.1038/s41564023-01593-7 (2024)

Kelly, C.L., Travis, N. M., Baya, P. A., Frey, C., Sun, X., Ward, B. B., Casciotti, K. L. Isotopomer labeling and oxygen dependence of hybrid nitrous oxide production. Biogeosciences, 21:3215–3238. DOI:10.5194/bg-21-3215-2024 (2024)

Wan, X. S., Sheng, H-X., Liu, L., Shen, H, Tang, W., Zou, W., Xu, M. N., Zheng, Z., Tan, E., Chen, M., Zhang, Y., Ward, B. B., Kao, S-J. Particle-associated denitrification is the primary source of N2O in oxic coastal waters. Nature Communications. 14: 8280, DOI:10.1038/s41467-023-43997-3 (2023)

Tang, T., Ward, B. B., Beman, M, Bristow, L., Clark, D., Fawcett, S., Frey. C, Fripiat, F., Herndl, G., Mdutyana, M., Paulot, F., Peng, X., Santoro, A., Shiozaki, T., Sintes, E., Stock, C., Sun, X., Wan, X., Xu, M., and Zhang, Y. Database of nitrification and nitrifiers in the global ocean. Earth System Science Data. 15: 5039–5077, DOI:10.5194/essd-15-5039-2023 (2023)

Tracey, J.C., Babbin, A.R., Wallace, E., Sun, X., Durussel, K.L., Frey, C., Martocello III, D.E., Tamasi, T. Oleynik, S. and Ward, B.B. All about nitrite: exploring nitrite sources and sinks in the eastern tropical North Pacific oxygen minimum zone. Biogeosciences. 20:12 pp. 2499-2523 DOI:10.5194/bg-20-2499-2023 (2023)

Wan, X.S., Hou, L., Kao, S.-J., Zhang, Y., Sheng, H.-X., Shen, H., Tong, S., Qin, W. and Ward, B.B., Pathways of N₂O production by marine ammonia-oxidizing archaea determined from dual-isotope labeling, Proceedings of the National Academy of Sciences of the United States of America, 120:11. DOI: 10.1073/pnas.2220697120 (2023)

Sun, X., Frey, C. and Ward, B. B., Nitrite oxidation across the full oxygen spectrum in the ocean. Global Biogeochemical Cycles. DOI: 10.1029/2022GB007548 (2023)

Frey, C. and Sun, X. and Szemberski, L. and Casciotti, K.L. and Garcia-Robledo, E. and Jayakumar, A. and Kelly, C.L. and Lehmann, M.F. and Ward, B.B., Kinetics of nitrous oxide production from ammonia oxidation in the Eastern Tropical North Pacific, Limnology and Oceanography, 68:2, 424-438, DOI: 10.1002/lno.12283 (2023)

Sheng, H.-X. Wan, X. S., Zou, B., Sun, Y., Hanoch, B., Zou, W., Yang, W., Wu, S., Huang, H., Ward, B. B. and Kao, S.-J. An efficient diazotroph derived nitrogen transfer pathway in coral reef system. Limnology and Oceanography, 16, pp. 29–36. DOI: 10.1002/lno.12324 (2023)

Wan, X. S., Hua-Xia Sheng, Dai, M., Casciotti, K. L., Church, M. J., Zou, W., Liu, L., Shen, H., Zhou. K., Ward, B. B. and Kao, S.-J. Epipelagic nitrous oxide production offsets carbon sequestration by the biological pump. Nature Geoscience. DIO: 10.1038/s41561-022-01090-2 (2022)

Kolody, B. C., Smith, S. R., Zeigler Allen, L., McCrow, J. P., Moustafa, A., Shi, D., Hopkinson, B. M., Morel, F. M. M., Ward, B. B. and Allen, A. E. Nitrogen and iron availability drive metabolic remodeling and natural selection of diverse phytoplankton during experimental upwelling.  mSystems, 7:5. DOI: 10.1128/msystems.00729-22 (2022)

Wan, X. S., Lin, H., Ward, B. B., Kao, S.-J. and Dai, M. Significant seasonal N₂O dynamics revealed by multi-year observations in the Northern South China Sea.  Global Biogeochemical Cycles, 6:10. DOI: 10.1029/2022GB007333 (2022)

Tang, W., Jayakumar, A., Sun, X., Tracey, J., Carroll, J. T., Wallace, E., Lee, J. A., Nathan, L. and Ward, B. B.  Nitrous oxide consumption in oxygenated and anoxic estuarine waters.  Journal of Geophysical Research.  DOI: 10.1029/2022GL100657 (2022)

Mdutyana, M., Marshall, T., Sun, X., Burger, J. M., Thomalla, S. J., Ward, B. B. and Fawcett, S. E. Controls on nitrite oxidation in the upper Southern Ocean: insights from winter kinetics experiments in the Indian sector Biogeosciences, 19:3425-3444. DIO: 10.5194/bg-19-3425-2022 (2022)

Tang, W., Tracey, J., Carroll, J. T., Wallace, E., Lee, J. A., Nathan, L., Sun, X., Jayakumar, A. and Ward, B. B.  Nitrous oxide production in Chesapeake Bay. Limnology and Oceanography, 67: 2101-2116. DOI: 10.1002/lno.12191 (2022)

Mdutyana, M., Sun, X, Burger, J. M., Flynn, R. F., Smith, S., van Horsten, N. R., Roychoudhury, A. N., Planquette, H., Bucciarelli, E., Thomalla, S. J., Ward, B. B., Fawcett, S. E. The kinetics of ammonium uptake and oxidation across the African sector of the Southern Ocean.  Limnology and Oceanography, 67: 973-991 DOI: 10.1002/lno.12050 (2022)

Carroll, J., Van Oostende, N., and Ward, B.B. Evaluation of Genomic Sequence-Based Growth Rate Methods for Synchronized Synechococcus Cultures. Applied and Environmental Microbiology, 88(1),e01743-21. DOI: 10.1128/AEM.01743-21 (2022)

Lueders-Dumont, J.A., Forden, A.G., Kast, E.R., Mohan, J.A., Walther, B.D., Sigman, D.M., and Ward, B.B. Controls on the nitrogen isotopic composition of fish otolith organic matter: Lessons from a controlled diet switch experiment. Geochimica et Cosmochimica Acta, 316, pp. 69-86. DOI: 10.1016/j.gca.2021.09.030 (2021)

Rees, A.P., Brown, I.J., Jayakumar, A., Lessin, G., Somerfield, P.J., and Ward, B.B. Biological nitrous oxide consumption in oxygenated waters of the high latitude Atlantic Ocean. Communications Earth and Environment, 2(1),36. DOI: s43247-021-00104-y (2021)

Wan, X.S. and Sheng, H.-X. and Dai, M. and Church, M.J. and Zou, W. and Li, X. and Hutchins, D.A. and Ward, B.B. and Kao, S.-J. Phytoplankton-Nitrifier Interactions Control the Geographic Distribution of Nitrite in the Upper Ocean. Global Biogeochemical Cycles, 35:11. DOI: 10.1029/2021GB007072 (2021)

Peng, X. and Ji, Q. and Angell, J.H. and Kearns, P.J. and Bowen, J.L. and Ward, B.B. Long-Term Fertilization Alters Nitrous Oxide Cycling Dynamics in Salt Marsh Sediments. Environmental Science and Technology, 55:15 10832-10842. DOI: 10.1021/acs.est.1c01542 (2021)

Sun, X. and Frey, C. and Garcia-Robledo, E. and Jayakumar, A. and Ward, B.B. Correction: Microbial niche differentiation explains nitrite oxidation in marine oxygen minimum zones. The ISME Journal, 15:5 1317-1329. DOI: 10.1038/s41396-021-01032-7 (2021) pdf

Schulz, K.G., Achterberg, E.P.,  Arístegui, J., Bach, L.T., Baños, I., Boxhammer, T., Erler, D., Igarza, M., Kalter, V., Ludwig, A., Löscher, C., Meyer, J., Meyer, J., Minutolo, F., Von Der Esch, E., Ward, B.B. and Riebesell, U. Nitrogen loss processes in response to upwelling in a Peruvian coastal setting dominated by denitrification - A mesocosm approach. Biogeosciences, 18:14 4305-4320. DOI: 10.5194/bg-18-4305-2021 (2021)

Sun, X. and Frey, C. and Garcia-Robledo, E. and Jayakumar, A. and Ward, B.B. Microbial niche differentiation explains nitrite oxidation in marine oxygen minimum zones. ISME Journal, 15:5 1317-1329, DOI: 10.1038/s41396-020-00852-3 (2021) pdf

Sun, X. and Jayakumar, A. and Tracey, J.C. and Wallace, E. and Kelly, C.L. and Casciotti, K.L. and Ward, B.B. Microbial N₂O consumption in and above marine N₂O production hotspots. ISME Journal, 15:5, 1434-1444.  DOI: 10.1038/s41396-020-00861-2 (2021) pdf

Bourbonnais, A., Frey, C., Sun, X., Bristow, L. A., Jayakumar, J., Ostrom, N. E., Casciotti, K. L. and Ward, B. B. Protocols for assessing transformation rates of nitrous oxide in the water column. Frontiers in Marine Science. DOI:10.3389/fmars.2021.611937 (2021) pdf

Sun, X., Frey, C., Garcia-Robledo, E., Jayakumar, A., and Ward, B. B. Microbial niche differentiation explains nitrite oxidation in marine oxygen minimum zones. ISME Journal, 15:1317-1329. (2021)

Sun, X., A. Jayakumar, J. Tracey, E. Wallace, C. Kelly, K. Casciotti, and B. B. Ward. Microbial N₂O consumption in and above marine N₂O production hotspots. ISME Journal, 15:1434-1444. DOI: 10.1038/s41396-020-00861-2  (2021).  pdf

Jayakumar, A., and Ward., B. B. Diversity and distribution of nitrogen fixation genes in the oxygen minimum zones of the world oceans. Biogeosciences, 17, 5953–5966. DOI:10.5194/bg-17-5953-2020  (2020)  pdf

Wilson, S. T., Al-Haj, A. N., Bourbonnais, A., Frey, C., Fulweiler, R. W., Kessler, J. D., Marchant, H. K., Milucka, J., Ray, N. E., Suntharalingham, P., Thornton, B. F., Upstill- Goddard, R. C., Weber, T. S., Arévalo-Martínez, D. L., Bange, H. W., Benway, H. M., Bianchi, D., Borges, A. V., Chang, B. X., Crill, P. M., del Valle, D. A., Farías, L., Joye, S. B., Kock, A., Labidi, J., Manning, C. C., Pohlman, J. W., Rehder, G., Sparrow, K. J., Tortell, P. D., Treude, T., Valentine, D. L., Ward, B. B., Yang, S. and Yurganov. L. N. Ideas and perspectives: A strategic assessment of methane and nitrous oxide measurements in the marine environment. Biogeosciences, 17, 5809–5828. DOI: 10.5194/bg-17-5809-2020 (2020) pdf

Mhlangabezi, M., Thomalla, S., Philibert, R., B. B. Ward and Fawcett, S. The seasonal cycle of nitrogen uptake and nitrification in the Atlantic sector of the Southern Ocean. Global Biogeochemical Cycles. DOI: 10.1029/2019GB006363 (2020) pdf

Lueders-Dumont, J. D., M. Sigman, B. Johnson, O. Jensen and B. B. Ward. Comparison of the isotopic composition of fish otolith-bound organic N with host tissue. Canadian Journal of Fisheries and Aquatic Sciences, 77: 264-275. DOI: 10.1139/cjfas-2018-0360 (2020) pdf

Lars Riis Damgaard, L. R., C. Kelly, K. L. Casciotti, B. B. Ward and N. P. Revsbech. Amperometric sensor for nanomolar nitrous oxide analysis. Analytica Chemica Acta., 1101: 135-140. DOI: 10.1016/j.aca.2019.12.019 (2020) pdf

Frey, C., H. W. Bange, E. P. Achterberg, A. Jayakumar, C. R. Löscher, D. L. Arévalo-Martínez, , E. León-Palmero, , M. Sun, X. Sun , R. C. Xie, S. Oleynik and B. B. Ward. Regulation of nitrous oxide production in low oxygen waters off the coast of Peru, Biogeosciences, 17:2263-2287. DOI: 10.5194/bg-2019-476 (2020) pdf

Rich, J. J., P. Arevalo, B. X. Chang, A.H. Devol, B. B. Ward. Anaerobic ammonium oxidation (anammox) and denitrification in Peru margin sediments. Journal of Marine Systems, 207. DOI: 10.1016/j.jmarsys.2018.09.007 (2020)

Babbin, A. R., C. Buchwald, F. M. M. Morel, S. D. Wankel, and B. B. Ward. Nitrite oxidation exceeds reduction and fixed nitrogen loss in anoxic Pacific waters. Marine Chemistry. DOI: 10.1016/j.marchem.2020.103814 (2020) pdf

Tracey, J., , M. Coronado, T. Giessen, M. VC. Y. Lau, P. Silver and B. B. Ward. The discovery of twenty-three new sequences encoding for encapsulin nanocompartments, including two in anammox bacteria. Scientific Reports, 9: 20122. DOI: 10.1038/s41598-019-56533-5 (2019)  pdf

Eveillard, D., N. Bouskill, D. Vintache, J. Gras, B. B. Ward and J. Bourdon. Probabilistic modeling of microbial networks for integrating partial quantitative knowledge within the nitrogen cycle. Frontiers in Microbiology, 9: 3298. DOI: 10.3389/fmicb.2918.03298 (2019) pdf

Chang, B. X., A. Jayakumar,  B. Widner, P. Bernhardt, C. W. Mordy, M. R. Mulholland and B. B. Ward. Low rates of dinitrogen fixation in the eastern tropical South Pacific. Limnology and Oceanography, 64: 1913-1923. DOI: 10.1002/lno.11159 (2019) pdf

Sun, X., Kop, L. F. M., Lau, M. C .Y., Frank, J., Jayakumar, A., Lücker, S. and Ward, B. B.. Uncultured Nitrospina-like species are major nitrite oxidizing bacteria in oxygen minimum zones. ISME Journal, 13:2391–2402. DOI: 10.1038/s41396-019-0443-7 (2019) pdf

Sabadel, A. J. M., , N. C. Van Oostende, B. B. Ward, , E. M. S. Woodward, R. Van Hale and , R. D. Frew. Characterization of particulate organic matter cycling during a summer North Atlantic phytoplankton bloom using amino acid C and N stable isotopes. Marine Chemistry, 214: 103670. DOI: 10.1016/j.marchem.2019.103670 (2019)  pdf

Van Oostende, N., R. Dussin, C. A. Stock, A. D. Barton, E. Curchitser, J. P. Dunne and B. B. Ward. Simulation of the ocean chlorophyll dynamic range from oligotrophy to coastal upwelling. Progress in Oceanography, 168: 232-247. DOI: 10.1016/j.pocean.2018.10.009 (2018) pdf

Jayakumar A., D. Balachandran, A. P. Rees, P. J. Kearns, J. L. Bowen and B. B. Ward. Community composition of nitrous oxide reducing bacteria investigated using a functional gene microarray. Deep-Sea Research, 156: 44-50. DOI: 10.1016/j.dsr2.2018.10.002 (2018) pdf

Ji, Q., E. Buitenhuis, P. Suntharalingam, J. L. Sarmiento and B. B. Ward. Global nitrous oxide production determined by oxygen sensitivity of nitrification and denitrification. Global Biogeochemical Cycles, 32: 1790-1802. DOI: 10.1029/2018GB005887 (2018) pdf

Ji, Q., C. Frey, X. Sin, Y.-S. Lee, A. Jayakumar, M. Jackson, J. C. Cornwell and B. B. Ward. Nitrogen and oxygen availabilities control water column nitrous oxide production during seasonal anoxia in the Chesapeake Bay. Biogeosciences, 15: 6127–6138. DOI: 10.5194/bg-15-6127-2018 (2018) pdf

Angell, J. H., A. Peng, Q. J, I. Craick, A. Jayakumar, P. J. Kearns, B. B. Ward and J. L. Bowen. Community composition of nitrous oxide related genes and their relationship to nitrogen cycling rates in salt marsh sediments. Frontiers in Microbiology, 9:170. DOI: 10.3389/fmicb.2018.00170 (2018) pdf

Lueders-Dumont, J., X. T. Wang, O. P. Jensen, D. M. Sigman and B. B. Ward. Nitrogen isotopic analysis of otolith-bound organic matter in modern and fossil fish otoliths. Geochimica et Cosmochimica Acta, 224:200-222. DOI: 10.1016/j.gca.2018.01.001 (2018) pdf

Peng, X., S. E. Fawcett, N. C. van Oostende, M. Wolf, D. Marconi, D. M. Sigman and B. B. Ward. Nitrogen assimilation and nitrification in the subarctic North Atlantic Ocean. Limnology and Oceanography, 63: 1462-1487. DOI: 10.1002/lno.10784 (2018) pdf

Jäntti, H., B. B. Ward, J. W. Dippner and S. Hietanen. Nitrification and the ammonia-oxidizing communities in the central Baltic Sea water column. Estuarine, Coastal and Shelf Science, 202: 280-289. DOI: 10.1016/j.ecss.2018.01.019  (2018) 

Marconi, D., D. M. Sigman, K. L. Casciotti, E. C. Campbell, M. A. Weigand, S. E. Fawcett, A. N. Knapp, B. B. Ward and G. H. Haug. Tropical dominance of nitrogen fixation in the North Atlantic Ocean. Global Biogeochemical Cycles, 31: 1608–1623. DOI: 10.1002/2016GB005613 (2017) pdf

Lisa, J. A., A. Jayakumar, B. B. Ward and B. Song. nirS denitrifying bacterial assemblages respond to environmental conditions of a shallow estuary. Environmental Microbiology Reports, 9: 766–778.  DOI: 10.1111/1758-2229.12594 (2017) pdf

Sun, X., Q. Ji, A. Jayakumar and B. B. Ward. Dependence of nitrite oxidation on nitrite and oxygen in low oxygen seawater. Geophysical Research Letters, 44: 7883-7891. DOI: 10.1002/2017GL074355 (2017) pdf

Jayakumar, A., B. X. Chang, B. Widner, P. Bernhardt, M. R. Mullholland and B. B. Ward. Biological nitrogen fixation in the oxygen minimum region of the Eastern Tropical North Pacific Ocean. ISME Journal, 11: 2356-2367. DOI: 10.1038/ismej.2017.97 (2017) pdf

Sun, X., A. Jayakumar and B. B. Ward. Community composition of nitrous oxide consuming bacteria in the oxygen minimum zone of the Eastern Tropical South Pacific. Frontiers of Microbiology, 8: 1183. DOI: 10.3389/fmicb.2017.01183 (2017) pdf

Ji, Q. and B. B. Ward. Nitrous oxide production in surface waters of the mid-latitude North Atlantic Ocean. JGR-Oceans, 122: 2612-2621. DOI: 10.1002/2016JC012467 (2017) pdf

Babbin, A. R., B. D. Peters, C. W. Mordy, B. Widner, K. L. Casciotti and B. B. Ward. Multiple metabolisms constrain the anaerobic nitrite budget in the Eastern Tropical South Pacific. Global Biogeochemical Cycles, 31: 258-271. DOI: 10.1002/2016GB005407 (2017) pdf

Van Oostende, N., S. E. Fawcett, D. Marconi, J. Lueders-Dumont, A. Sabadel, W. M. S. Woodward, B. Jonsson, D. M. Sigman and B. B. Ward. Variation of summer phytoplankton community composition and its relationship to nitrate and ammonium assimilation across the North Atlantic Ocean. Deep-Sea Research I, 121: 79-94. DOI: 10.1016/j.dsr.2016.12.012 (2017) pdf

Peters, B. D., A. R. Babbin, B. B. Ward, K. A. Lettmann, C. W. Mordy, O. Ulloa and K. L. Casciotti. Vertical modeling of the nitrogen cycle in the eastern tropical south Pacific oxygen deficient zone using high resolution concentration and isotope measurements. Global Biogeochemical Cycles, 30: 1661-1681. DOI: 10.1002/2016GB005415 (2016) pdf

Horak, R. E. A., W. Reuf, B. B. Ward and A.H. Devol. Expansion of denitrification and anoxia in the eastern tropical North Pacific from 1972 to 2012. Geophysical Research Letters, 43(10): 5252-5260. DOI: 10.1002/2016GL068871 (2016) pdf

Peng, X., C. A. Fuchsman, A. Jayakumar, M. J. Warner, A. H. Devol and B. B. Ward. Revisiting nitrification in the Eastern Tropical South Pacific: A focus on controls. J. Geophys. Res. Oceans, 121. DOI:10.1002/2015JC011455 (2016) pdf

Peng, Xuefeng, Qixing Ji, Angell J. H, Kearns, P. J., Yang, H. J., Bowen, J. L. and B. B. Ward. Long-term fertilization alters the relative importance of nitrate reduction pathways in salt marsh sediments. J. Geophys. Res. Biogeosci., 121. DOI:10.1002/2016JG003484 (2016) pdf

Rees, A. P., I. J. Brown, A. Jayakumar and B. B. Ward. The inhibition of N2O production by ocean acidification in cold temperate and polar waters. Deep-Sea Research Part II, 127:93-101. DOI: 10.1016/j.dsr2.2015.12.006 (2016) pdf

Ward, B. B. and N. Van Oostende. Phytoplankton assemblage during the North Atlantic spring bloom assessed from functional gene analysis. J. Plankton Res. 00(00):1-16. DOI:10.1093/plankt/fbw043 (2016) pdf

Ji, Q., A. R. Babbin, X. Peng, J. Bowen and B. B. Ward. Nitrogen substrate–dependent nitrous oxide cycling in salt marsh sediments. Journal of Marine Research, 73:71-92. DOI: 10.1357/002224015815848820 (2015) pdf

Ji., Q., A. R. Babbin, A. Jayakumar, S. Oleynik and B. B. Ward.  Nitrous oxide production by nitrification and denitrification in the Eastern Tropical South Pacific oxygen minimum zone. Geophys. Res. Let. 42:24, p.p. 10,755–10,764. DOI:10.1002/2015GL066853 (2015) pdf

Rees, A. P., I. J. Brown, A. Jayakumar and B. B. Ward. The inhibition of N₂O production by ocean acidification in cold temperate and polar waters. Deep Sea Research Part II: Topical Studies in Oceanography 127, p.p. 93-101. DOI:10.1016/j.dsr2.2015.12.006 (2015) pdf

Babbin, A. R., A. Jayakumar and B. B. Ward. Organic Matter Loading Modifies the Microbial Community Responsible for Nitrogen Loss in Estuarine Sediments. Microb. Ecol., 71:3, p.p. 555–565. DOI:10.1007/s00248-015-0693-5 (2015) pdf

Zhang, E., S. Huang, Q. Ji, M. Silvernagel, Y. Wang, B. B. Ward, D. Sigman and G. Wysocki. Nitric Oxide Isotopic Analyzer Based on a Compact Dual-Modulation Faraday Rotation Spectrometer. Sensors25992-26008. DOI:10.3390/s151025992 (2015) pdf

Babbin, A. R., D. Bianchi, A. Jayakumar and B. B. Ward. Rapid nitrous oxide cycling in the suboxic ocean. Science 348:6239, pp. 1127-1129. DOI:10.1126/science.aaa8380 (2015) pdf

Fawcetta, S. F., B. B. Ward, M. W. Lomasb and D. M. Sigman. Vertical decoupling of nitrate assimilation and nitrification in the Sargasso Sea. Deep-Sea Research I 103:64–72. DOI:10.1016/j.dsr.2015.05.004 (2015) pdf

Bowen, J. L., D. Weisman, M. Yasuda, A. Jayakumar, H. G. Morrison and B. B. Ward. Marine Oxygen-Deficient Zones Harbor Depauperate Denitrifying Communities Compared to Novel Genetic Diversity in Coastal Sediments. Microb Ecol 70:311–321. DOI:10.1007/s00248-015-0582-y (2015) pdf

Van Oostende, N., J. P. Dunne, S. E. Fawcett and B. B.Ward. Phytoplankton succession explains size-partitioning of new production following upwelling-induced blooms. Journal of Marine Systems 148:14–25.  DOI:10.1016/j.jmarsys.2015.01.009 (2015) pdf 

Peng, X., C. A. Fuchsman, A. Jayakumar, M. J. Warner, A. H. Devol and  B. B. Ward. Revisiting nitrification in the Eastern Tropical South Pacific: A focus on controls.  Journal of Geophysical Research: Oceans 121. DOI: 10.1002/2015JC011455 (2016) pdf   Data file

Peng, X., C. A. Fuchsman, A. Jayakumar, S. Oleynik, W. Martens-Habbena, A. H. Devol and B. B. Ward. Ammonia and nitrite oxidation in the Eastern Tropical North Pacific. Global Biogeochemical Cycles 29. DOI:10.1002/2015GB005278 (2015) pdf  Data file

Tiano, L., E. G. Robledo, T. Dalsgaard, A. H. Devol, B. B. Ward, O. Ulloa, D. E. Canfield and N. P. Revsbech.  Oxygen distribution and aerobic respiration in the north and south eastern tropical Pacific oxygen minimum zones.  Deep Sea Research I 194: 173-183. DOI: 10.1016/j.dsr.2014.10.001 (2014) pdf

Bowen, J. L., A. R. Babbin, P. J. Kearns and B. B. Ward. Connecting the dots:  Linking nitrogen cycle gene expression to nitrogen fluxes from marine sediment mesocosms.  Frontiers in Microbiology 5:429.  DOI: 10.3389/fmicb.2014.00429  (2014) pdf

Tait, K., V. Kitidis, B. B. Ward, D. G. Cummings, M. R. Jones, P. J. Somerfield and S. Widdicombe.  Spatio-tempral variability in ammonia oxidation and ammonia oxidising bacteria and archaea in coastal sediments of the Western English Channel. Marine Ecology Progress Series 511:41-58. DOI: 10.3354/meps10933 (2014) pdf

Shilova, I. N., J. C. Robidart, H. J. Tripp, K. Turk-Kubo, B. Wawrik, A. F. Post, A. W. Thompson, B. B. Ward, J. T. Hollibaugh, A. Millard, M. Ostrowski, D. Scanlan, R. W. Paerl, R. Stuart and J. P. A. Zehr. A microarray for assessing gene transcription from pelagic marine microbial taxa.  ISME-J 8: 1476-1491. DOI: 10.1038/ismej.2014.1 (2014) pdf

Chang, B. X., J. R. Rich, A. Jayakumar, H. Naik, A. Prathihary, R. G. Keil, B. B. Ward and A. H. Devol.  The effect of organic carbon on nitrogen loss in the oxygen deficient waters of the Eastern Tropical Pacific and Arabian Sea. Limnology and Oceanography 59: 1267-1274. DOI: 10.4319/lo.2014.59.4.1267 (2014) pdf

Babbin, A.R., R. G. Keil, A. H. Devol and B. B. Ward. Organic Matter Stoichiometry, Flux and Oxygen Control Nitrogen Loss in the Ocean. Science , 344:406-408. DOI: 10.1126/science.1248364 (2014) pdf 

Fawcett, S.E., M. W. Lomas, B. B. Ward and D. M. Sigman. The counter intuitive effect of summer-to-fall mixed layer deepening on eukaryotic new production in the Sargasso Sea.  Global Biogeochemical Cycles 28(2):86-102.  DOI: 10.1002/2013GB004579 (2014) pdf 

Ward, B. B. Nitrification. In Earth Systems and Environmental Sciences.  Elsevier. DOI: 10.1016/B978-0-12-409548-9.00697-7 (2013) pdf

Jayakumar, A., X. Peng and B. B. Ward. Community composition of ammonia-oxidizing archaea from surface and anoxic depths of oceanic oxygen minimum zones.  Frontiers of Aquatic Microbiology 4: Article 177. DOI: 10.3389/fmicb.2013.00177 (2013) pdf

Bowen, J. L., P. J. Kearns, M. Holcomb and B. B. Ward. Acidification alters the community composition of ammonia oxidizing microbial assemblages in marine mesocosms. Marine Ecology Progress Series 492: 1-8. DOI: 10.3354/meps10526 (2013) pdf

Francis, C. A., G. D. O’Mullan, J. C. Cornwell and B. B. Ward. Transitions in nirS-type denitrifier diversity, community composition and biogeochemical activity along the Chesapeake Bay Estuary. Frontiers of Microbiology. DOI: 10.3389/fmicb.2013.00237 (2013) pdf

Ward, B. B. and M. M. Jensen. The microbial nitrogen cycle. Frontiers in Microbiology, 5, 1-2. [553]. DOI: 10.3389/fmicb.2014.00553 (2014) pdf

Ward, B. B. How Nitrogen is Lost.  Science 341:352-353. DOI: 10.1126/science.1240314 (2013) pdf

Peng, X., A. Jayakumar and B. B. Ward. Community composition of ammonia-oxidizing archaea from surface and anoxic depths of oceanic oxygen minimum zones.  Frontiers of Microbiology.  DOI: 10.3389/fmicb.2013.00177  (2013) pdf

Newell, S. E., S. E. Fawcett and B. B. Ward. Depth distribution of ammonia oxidation rates and ammonia-oxidizer community composition in the Sargasso Sea.  Limnology and Oceanography 58:1491-1500. DOI: 10.4319/lo.2013.58.4.1491 (2013) pdf

Babbin, A. R and B. B. Ward. Controls on nitrogen loss processes in Chesapeake Bay sediments.  Environmental Science and Technology. DOI: 10.1021/es304842r (2013) pdf

Voss, M., H. W. Bange, J. W. Dippner, J. J. Middelburg, J. P. Montoya and B. B. Ward. The marine nitrogen cycle:  Recent discoveries, uncertainties and the potential relevance of climate change.  Philosophical Transactions of the Royal Society B.  368: 20130121. DOI: 10.1098/rstb.2013.0121 (2013) pdf

Ward, B. B.  The Global Nitrogen Cycle.  In: A. H. Knoll, D. E. Canfield and K. O. Konhauser, Editors, Fundamentals of Geomicrobiology, Wiley-Blackwell, Chichester, UK, Pp. 36-48. DOI: 10.1002/9781118280874.ch4  (2012) pdf

Jayakumar, A., M. M. D. Al-Rashaidat, B. B. Ward and M. R. Mulholland. Diversity, distribution and expression of nifH genes in oxygen deficient waters of the Arabian Sea. FEMS Microbial Ecology. DOI: 10.1111/j.1574-6941.2012.01430.x (2012) pdf

Kritee, K., D. M. Sigman, J. Granger, B. B. Ward, A. Jayakumar and C. Deutsch. Reduced isotope fractionation by denitrification under conditions relevant to the ocean. Geochemica Cosmochemica Acta, 92:243-259. DOI: 10.1016/j.gca.2012.05.020  (2012) pdf

Moffett, J., C. B. Tuit and B. B. Ward. Chelator-induced inhibition of copper metalloenzymes in denitrifying bacteria. Limnology and Oceanography, 57:272-280. DOI:10.4319/lo.2012.57.1.0272 (2012) pdf

Bouskill, N. J., D. Eveillard, D. M. Chien, A. Jayakumar and B. B. Ward, Environmental factors determining ammonia‐oxidizing organism distribution and diversity in marine environments. Environmental Microbiology 14:714-729. DOI: 10.1111/j.1462-2920.2011.02623.x (2012) pdf

Newell, S.E., Babbin, A. R., Jayakumar, A. and B. B. Ward. Ammonia oxidation rates and nitrification in the Arabian Sea. Global Biogeochemical Cycles, 25 GB4016. DOI:10.1029/2010GB003940 (2011) pdf

Bouskill, N. J., D. Eveillard, D. M. Chien, A. Jayakumar and B. B. Ward, Environmental factors determining ammonia-oxidizing organism distribution and diversity in marine environments. Environmental Microbiology. DOI: 10.1111/j.1462-2920.2011.02623.x (2011) pdf

Fawcett, S. E., M. W. Lomas, J. R. Casey, B. B. Ward and D. M. Sigman. Assimilation of upwelled nitrate by small eukaryotes in the Sargasso Sea. Nature Geosciences, 4: 717-722.  DOI: 10.1038/ngeo1265 (2011) pdf

Bhadury, P. and Ward, B. B. Intron features of key functional genes mediating nitrogen metabolism in marine phytoplankton. Marine Genomics, 3: 207-213. DOI: 10.1016/j.margen.2011.06.002 (2011) pdf

Ward,B. B., A. P. Rees, P. J. Somerfield and I. R. Joint. Linking phytoplankton community composition to seasonal changes in f ratio. ISME Journal, 5 : 1759-1770. DOI:10.1038/ismej.2011.50 (2011) pdf

Bowen, J. L., B. B. Ward, H. G. Morrison, J. E. Hobbie, I. Valiela, L. A. Deegan and M. L. Sogin. Microbial community composition in sediments resists perturbation by nutrient enrichment. ISME Journal, 5 : 1540-1548. DOI:10.1038/ismej.2011.22 (2011) pdf

Fawcett, S. E. and B. B. Ward. Phytoplankton succession and nitrogen utilization during the development of a simulated upwelling bloom. Marine Ecology Progress Series, 436: 13-31.  DOI: 10.3354/meps09070 (2011) pdf

Ward, B. B. Nitrification in the Ocean. In: B. B. Ward, M. G. Klotz and D. A. Arp, Editors. Nitrification, ASM Press, Washington, D.C. 325-345. DOI: 10.1128/9781555817145 (2011) pdf

Ward, B. B. Measurement and distribution of nitrification rates in the oceans. In Microbial Nitrification and Related Processes. M. G. Klotz, editor, Methods in Enzymology, 486: 307-323. DOI: 10.1016/B978-0-12-381294-0.00013-4 (2011) pdf

Ward, B. B. and N. J. Bouskill. The utility of functional gene arrays for assessing community composition, relative abundance and distribution of ammonia-oxidizing bacteria and archaea.  In Microbial Nitrification and Related Processes. M. G. Klotz, editor. Methods in Enzymology, 496: 373-396. DOI:10.1016/B978-0-12-386489-5.00015-4 (2011) pdf

Campbell, M., P. Chain, H. Dang, A. El Sheik, J. Norton, N. Ward, B. B. Ward and M. G. Klotz. Nitrosococcus watsonii sp. nov., a new species of marine obligate ammonia-oxidizing bacteria that is not omnipresent in the world’s oceans. FEMS Microbiology Ecology, 76: 39-48. DOI:10.1111/j.1574-6941.2010.01027.x (2011) pdf

Bouskill, N. J., D. Eveillard, G. D. O’Mullan, G. A. Jackson and B. B. Ward. Seasonal and annual reoccurrence in Ammonia-oxidizing bacterial populationstructure. Environmental Microbiology 13: 872-886. DOI:10.1111/j.1462-2920.2010.02362.x (2011) pdf

Malcolm, E. G., J. K. Schaefer, E. B. Ekstrom, C. B. Tuit, A. Jayakumar, H. Park, B. B. Ward and F. M. M. Morel. Mercury methylation in oxygen deficient zones of the oceans: No evidence for the predominance of anaerobes. Marine Chemistry, 122: 11-19. DOI:10.1016/j.marchem.2010.08.004 (2010) pdf

Bulow, S.E., J. J. Rich, H. Naik, A. Pratihary. and B. B. Ward. Denitrification and not anammox is dominant in the Arabian Sea Oxygen Minimum Zone. Deep-Sea Res. Pt 1, 57: 384-393. DOI:10.1038/nature08276 (2010) pdf

Bhadury, P. and B. B. Ward. Molecular diversity of marine phytoplankton communities based on key functional genes. Journal of Phycology 45: 1335-1347. DOI:10.1111/j.1529-8817.2009.00766.x (2009) pdf

Jayakumar, A., G. O’Mullan, S. W. A. Naqvi and B. B. Ward Distribution and Relative Quantification of key Genes Involved in Fixed Nitrogen Loss From the Arabian Sea Oxygen Minimum Zone. In, Indian Ocean Biogeochemical Processes and Ecological Variability eds., Wiggert J.D., Hood R.R., Naqvi, S.W.A., Brink, K.H. and Smith, S.L.  American Geophysical Union, 187-203. DOI:10.1029/2008GM000730 (2009) pdf

Ward, B. B., A. H. Devol, J. J. Rich, B. X. Chang, S. E. Bulow, H. Naik, A. Pratihary and A. Jayakumar. Denitrification as the dominant nitrogen loss process in the Arabian Sea. Nature 461: 78-82. DOI:10.1038/nature08276 (2009) pdf

Jayakumar, A., G. O’Mullan, S. W. A. Naqvi and B. B. Ward. Denitrifying bacterial community composition changes associated with stages of denitrification in oxygen minimum zones. DOI:10.1007/s00248-009-9487-y Microb. Ecol. 58, 350-362. (2009) pdf

Song, B. K., E. Chyun, P. Jaffe and B. B. Ward. Molecular methods to detect and monitor of uncultured dissimilatory arsenate respiring bacteria (DARB) in sediments. FEMS Microbial Ecology, 68: 108-117. DOI:10.1111/j.1574-6941.2009.00657.x (2009) pdf

Ward, B. B., C.B Tuit, A. Jayakumar, J.J. Rich, J. Moffett and W. Naqvi. Organic carbon and not copper, controls denitrification in oxygen minimum zones of the ocean. Deep Sea Research I 55: 1672-1683. DOI:10.1016/j.dsr.2008.07.005 (2008) pdf

Ward, B. B. Phytoplankton community composition and gene expression of functional genes involved in carbon and nitrogen assimilation. Journal of Phycology 44: 1490-1503. DOI:10.1111/j.1529-8817.2008.00594.x (2008)  pdf

Ward, B. B. Nitrification. In: Nitrogen in the Marine Environment. Eds. D. G. Capone, D. A. Bronk, M. R. Mulholland and E. J. Carpenter. Elsevier, Amsterdam. 199-262. ISBN: 9780123725226 (2008) pdf

Bulow, S. E., C. A. Francis, G. A. Jackson and B. B. Ward. Sediment denitrifier community composition and nirS gene expression investigated with functional gene microarrays. Environmental Microbiology 10: 3057-3069. DOI:10.1111/j.1462-2920.2008.01765.x (2008) pdf

Starkenburg, S. R., F. W. Larimer, L. Y. Stein, M. G. Klotz, P. S. Chain, L. A. Sayavedra-Soto, A. T. Poret-Peterson, M. E. Gentry, D. J. Arp, B. B. Ward and P. J. Bottomley. Complete genome sequence of Nitrobacter hamburgensis X14 and comparative genomic analysis of species within the genus Nitrobacter.  Appl Environ Microbiol 74: 2852-2863. DOI: 10.1128/AEM.02311-07 (2008) pdf

Ward, B. B. Nitrification. In: Encyclopedia of Ecology. Eds. S. E. Jorgensen and B. D. Faith, Ecological Processes. Vol 3 of Encyclopedia of Ecology, 5 vols. Elsevier, Oxford. Pp. 2511-2518. DOI:10.1016/B978-008045405-4.00280-9 (2008) pdf

Duce, R.A., J. LaRoche, K. Altieri, K. Arrigo, A. Baker, D.G. Capone, S. Cornell, F. Dentener, J. Galloway, R.S. Ganeshram, R.Geider, T. Jickells, M.M. Kuypers, R. Langlois, P. S. Liss, S. M. Liu, J.J. Middelburg, C.M. Moore, S. Nickovic, A. Oschlies, T. Pedersen, J. Prospero, R. Schlitzer, S. Seitzinger, L.L. Sorensen, M. Uematsu, O. Ulloa, M. Voss, B. B. Ward and L. Zamora. Impacts of Atmospheric Anthropogenic Nitrogen on the Open Ocean. Science, 320:893-898. DOI: 10.1126/science.1150369 (2008) pdf

Rich, J. J., O. R. Dale, B. K. Song and B. B. Ward. Anaerobic ammonium oxidation (anammox) in Chesapeake Bay sediments. Microbial Ecology, 55: 311-320. DOI: 10.1007/s00248-007-9277-3 (2008) pdf

Ward, B. B. Nitrogen cycling in aquatic environments. In: Manual of Environmental Microbiology. Eds. C. J. Hurst, R. L. Crawford, J. L. Garland, D. A. Lipson, A. L. Mills, L. D. Stetzenbach. American Society for Microbiology, New York. 511-522. DOI: 10.1128/9781555815882.ch42  (2007) 

Adhitya, A. , F. I. M. Thomas and B. B. Ward. Diversity of assimilatory nitrate reductase genes from plankton and epiphytes associated with a seagrass bed. Microbial Ecology, 54: 587-597. DOI: 10.1007/s00248-006-9175-0 (2007) pdf

Ward, B. B., D. Eveillard, J. D. Kirshtein, J. D. Nelson, M. A. Voytek and G. A. Jackson. Ammonia-oxidizing bacterial community composition in estuarine and oceanic environments assessed using a functional gene microarray. Environmental Microbiology 9: 2522-2538. DOI: 10.1111/j.1462-2920.2007.01371.x (2007) pdf

Song, B. K. and B. B. Ward.  Molecular cloning and characterization of high affinity nitrate transporters in marine phytoplankton. Journal of Phycology 43:542-552. DOI: 10.1111/j.1529-8817.2007.00352.x (2007) pdf

Moisander, P. H., A. E. Morrison, B. B. Ward, B. D. Jenkins and J. P. Zehr. Spatial-temporal variability in diazotroph assemblages in Chesapeake Bay using an oligonucleotide nifH microarray. Environmental Microbiology, 9:1823-1835. DOI: 10.1111/j.1462-2920.2007.01304.x  (2007) pdf

Taroncher-Oldenburg, G. and B. B. Ward. Oligonucletoide microarrays for the study of microbial communities. In: DNA Analysis by Nonradioactive Probes (Ed.) Hilario, E. and Mackay, J.F., Humana Press, Totowa, NY, USA pp 301-315. DOI: 10.1385/1-59745-229-7:301 (2006) pdf

Klotz, M. G., D. J. Arp, P. S. G. Chain, A. R. El-Sheikh, L. J. Hauser, N. G. Hommes, F. W. Larimer, S. A. Malfatti, J. M. Norton, A. T. Poret-Peterson, L. M. Vergez and B. B. Ward. Complete genome sequence of the marine, chemolithoautotrophic ammonia-oxidizing bacterium Nitrosococcus oceani ATCC 19707. Applied and Environmental Microbiology, 72: 6299-6315. DOI: 10.1128/AEM.00463-06 (2006) pdf

Glatz, R. E., P. W. Lepp, B. B. Ward and C. A. Francis. Planktonic microbial community composition across steep physical/chemical gradients in permanently ice‐covered Lake Bonney, Antarctica. Geobiology, 4: 53-67.  DOI: 10.1111/j.1472-4669.2006.00057.x (2006) pdf

Bronk, D. A. and B. B. Ward. Inorganic and organic nitrogen cycling in the Southern California Bight. (Deep-Sea Research, 52: 2285-2300.  DOI: 10.1016/j.dsr.2005.08.002  (2005) pdf

Ward, B. B. and G. D. O’Mullan. Community level analysis: Genetic and biogeochemcial approaches to investigate community composition and function in aerobic ammonia oxidation. In: Methods in Enzymology, 397:395-413.  DOI: 10.1016/S0076-6879(05)97024-9 (2005)  pdf

Ward, B. B. Temporal variability in nitrification rates and related biogeochemical factors in Monterey Bay, California. Marine Ecology-Progress Series, 292: 97-109. DOI: 10.3354/meps292097  (2005) pdf

Ward, B. B. Molecular approaches to marine microbial ecology and the marine nitrogen cycle. In: Annual Review of Earth and Planetary Science, 33:092203.122514. DOI: 10.1146/annurev.earth.33.092203.122514 (2005)  pdf

Song, B. and B. B. Ward. Genetic diversity of benzoyl coenzyme A reductase genes detected in denitrifying isolates and estuarine sediment communities. Applied and Environmental Microbiology, 71: 2036-2045. DOI: 10.1128/AEM.71.4.2036-2045.2005 (2005) pdf

Ward, B. B., J. Granger, M. T. Maldonado, K. L. Casciotti, S. Harris and M. L. Wells. Denitrification in the hypolimnion of permanently ice-covered Lake Boney, Antarctica. Aquatic Microbial Ecology, 52: 197-205. DOI: 10.3354/ame038295 (2005) pdf

Casciotti, K. L. and B. B. Ward. Phylogenetic analysis of nitric oxide reductase gene homologues from aerobic ammonia-oxidizing bacteria. FEMS Microbial Ecology, 52: 197-205.  DOI: 10.1016/j.femsec.2004.11.002  (2005) pdf

Allen, A. E., B. Song and B. B. Ward. Characterization of diatom (Bacillariophyceae) nitrate reductase genes and detection of eukaryotic nitrate reductase genes from marine waters. Journal of Phycology, 41: 95-104.  DOI: 10.1111/j.1529-8817.2005.04090.x  (2005)  pdf

O’Mullan, G. D. and B. B. Ward. Relationship of Temporal and Spatial Variabilities of Ammonia-Oxidizing Bacteria to Nitrification Rates in Monterey Bay, CA. Applied and Environmental Microbiology, 71: 697-705. DOI: 10.1128/AEM.71.2.697-705.2005  (2005) pdf

Jiang, W., A. Saxena, B. Song, B. B. Ward, T. J. Beveridge and S. C. B. Myneni. Elucidation of functional groups on Gram-positive and Gram-negative bacterial surfaces using infrared spectroscopy, Langmuir, 20 11433-11442.  DOI: 10.1021/la049043+ (2004) pdf

Song, B. and B. B. Ward. Molecular characterization of the assimilatory nitrate reductase gene and its expression in the marine green alga Dunaliella tertiolecta. Journal of Phycology, 40: 721-731.  DOI: 10.1111/j.1529-8817.2004.03078.x  (2004) pdf

Jenkins, B. D., G. F. Steward, S. M. Short, B. B. Ward and J. P. Zehr. Fingerprinting diazotroph communities in the Chesapeake Bay by using a DNA macroarray. Applied and Environmental Microbiology, 70: 1767-1776. DOI: 10.1128/AEM.70.3.1767-1776.2004  (2004). pdf

Steward, G. F., B. D. Jenkins, B. B. Ward and J. P. Zehr. Development and testing of a DNA macroarray to assess nitrogenase (nifH) gene diversity. Applied and Environmental Microbiology, 70: 1455-1465  DOI: 10.1128/AEM.70.3.1455-1465.2004  (2004) pdf

Jayakumar, D. A., C. A. Francis, S. W. A. Naqvi and B. B. Ward. Diversity of nitrite reductase genes in the denitrifying water column of the coastal Arabian Sea. Aquatic Microbial Ecology, 34: 69-78. DOI: 10.3354/ame034069  (2004) pdf

Francis, C. A., G. D. O’Mullan and B. B. Ward. Diversity of ammonia monooxygenase (amoA) genes across environmental gradients in Chesapeake Bay sediments. Geobiology, 1: 129-140.  DOI: 10.1046/j.1472-4669.2003.00010.x  (2003) pdf

Ward, B. B.  Significance of anaerobic ammonium oxidation in the ocean. Trends in Microbiology, 11: 408-410.  DOI: 10.1016/S0966-842X(03)00181-1 (2003) pdf

Casciotti, K. L., D. M. Sigman and B. B. Ward. Linking Diversity and Stable Isotope Fractionation in Ammonia-Oxidizing Bacteria. Geomicrobiology Journal, 20: 335-353. DOI: 10.1080/01490450303895  (2003) pdf

Taroncher-Oldenburg, G, E. Griner, C. A. Francis and B. B. Ward. Oligonucleotide microarray for the study of functional gene diversity of the nitrogen cycle in the environment. Applied and Environmental Microbiology, 69: 1159-1171. DOI: 10.1128/AEM.69.2.1159-1171.2003  (2003) pdf

Ward, B. B., J. Granger, M. T. Maldonado and M. L. Wells. What limits bacterial production in the suboxic region of permanently ice-covered Lake Bonney, Antarctica? Aquatic Microbial Ecology, 31: 33-47. DOI: 10.3354/ame031033  (2003) pdf

Caffrey, J. M., N. E. Harrington, I. P. Solem and B. B. Ward. Biogeochemical Processes in a Small California Estuary, Elkhorn Slough, CA.: 2. Nitrification Activity, Community Structure and Role in Nitrogen Budgets, Marine Ecology Progress Series, 248: 27-40.  DOI: 10.3354/meps233039  (2003) pdf

Song, B. and B. B. Ward. Nitrite reductase genes in halobenzoate degrading denitrifyng bacteria and related species. FEMS Microbial Ecology, 34: 349-357.  DOI: 10.1111/j.1574-6941.2003.tb01075.x  (2003) pdf

Granger, J. and B. B. Ward. Accumulation of nitrogen oxides in copper-limited cultures of denitrifying bacteria. Limnology and Oceanography, 48: 313-318. DOI: 10.4319/lo.2003.48.1.0313 (2003) pdf

Ward, B. B. and G. D. O'Mullan. Worldwide distribution of marine ammonia-oxidizing Gamma-Proteobacteria detected in seawater by PCR and sequencing of 16S rRNA and AmoA genes. Applied and Environmental Microbiology, 68: 4153-4157.  DOI: 10.1128/AEM.68.8.4153-4157.2002  (2002) pdf

Ward, B. B. How many species of prokaryotes are there? Proceedings of the National Academy of Sciences, US. 99:10234-10236. DOI: 10.1073/pnas.162359199  (2002) pdf

Caffrey, J. M., N. E. Harrington and B. B. Ward. Biogeochemical Processes in a Small California Estuary: 1. Benthic Fluxes and Pore Water Constituents Reflect High Nutrient Freshwater Inputs. Marine Ecology Progress Series, 233: 39-53. DOI: 10.3354/meps233039 (2002) pdf

Ward, B. B. Nitrification in Aquatic Systems. Encyclopedia of Environmental Microbiology. D. A Capone, Ed., Wiley & Sons, New York, Pp. 2144-2167. DOI: 10.1002/0471263397.env287 (2002)

Zehr, J. P and B. B. Ward. Nitrogen cycling in the ocean: new perspectives on processes and paradigms. Applied and Environmental Microbiology, 68: 1015-1024.  DOI: 10.1128/AEM.68.3.1015-1024.2002 (2002) pdf

Ward, B. B. and D. A. Bronk. Net nitrogen uptake and DON release in surface waters: Size fraction experiments implicate grazing and community structure in DON release. Marine Ecology Progress Series, 219: 11-24. (2001) pdf

Golet, D. S. and B. B. Ward. Vertical distribution of denitrification potential, denitrifying bacteria and benzoate utilization in intertidal microbial mat communities. Microbial Ecology, 42: 22-34. DOI: 10.1007/s002489900184 (2001)

Casciotti, K. A. and B. B. Ward. Nitrite reductase genes in ammonia-oxidizing bacteria. Applied and Environmental Microbiology, 67: 2213-2221. DOI: 10.1128/AEM.67.5.2213-2221.2001 (2001) pdf

Bothe, H., G. Jost, M. Schloter, B. B. Ward and K.-P. Witzel. Molecular analysis of ammonia oxidation and denitrification in natural environments. DOI: 10.1111/j.1574-6976.2000.tb00566.x  FEMS Microbiological Reviews, 24: 673-690. (2000) pdf

Ward, B. B. Nitrification and the marine nitrogen cycle. In: D. Kirchman, ed. Microbial Ecology of the Oceans, Wiley-Liss, New York, pp 427-454. ISBN-10: 0471299928 (2000)  pdf

Bronk, D. A. and B. B. Ward. Magnitude of DON release relative to gross nitrogen uptake in marine systems. Limnology and Oceanography, 45: 1879-1883. DOI: 10.4319/lo.2000.45.8.1879 (2000) pdf

Ward, B. B., D. P. Martino, M. C. Diaz and S. B. Joye. Analysis of ammonia-oxidizing bacteria from hypersaline Mono Lake, California on the basis of 16s rRNA sequences. Applied and Environmental Microbiology, 67: 2873-2881. DOI: 10.1128/aem.66.7.2873-2881.2000 (2000) pdf

Bronk, D. A. and B. B. Ward. Gross and net nitrogen uptake and DON release in the euphotic zone of Monterey Bay, California. Limnology and Oceanography, 44:573-585. DOI: 10.4319/lo.1999.44.3.0573 (1999) pdf

Voytek, M. A., B. B. Ward and J. C. Priscu. The distribution and relative abundance of ammonia-oxidizing bacteria in six Antarctic Dry Valley lakes. Hydrobiologia, 401: 113-130. DOI: 10.1023/A:1003754830988 (1999) pdf

Francis, C. A., A. K. Francis, D. S. Golet and B. B. Ward. Quantification of catechol 2,3-dioxygenase gene homology abundance in intertidal sediments. Aquatic Microbial Ecology, 15: 225-231. DOI: 10.3354/ame015225 (1998)

Hogan, M. E. and B. B. Ward. Response of a marine sediment microbial community exposed to 2,4-dichlorophenoxyacetic acid. Microbial Ecology, 35: 72-82. DOI: 10.1007/s002489900061 (1998) pdf

Voytek, M. A., B. B. Ward and J. C. Priscu. The abundance of ammonia-oxidizing bacteria in Lake Bonney, Antarctica determined by immunofluorescence, PCR and in situ hybridization. Antarctic Research Series, The McMurdo Dry Valleys, pp. 217-228. DOI: 10.1029/AR072p0217 (1998)

Diaz, C. M. and B. B. Ward. Sponge mediated nitrification in tropical benthic communities. Marine Ecology Progress Series, 156: 97-107. DOI: 10.3354/meps156097 (1997)

Bard, D. G. and B. B. Ward. A species-specific bacterial productivity method using immunomagnetic separation and radiotracer experiments. Journal of Microbiological Methods, 28: 207-219. DOI: 10.1016/S0167-7012(97)00987-1 (1997) pdf

Ward, B. B. and J.C. Priscu. Detection and characterization of denitrifying bacteria in an ice-covered Antarctic Lake. Hydrobiologia, 347: 57-68. DOI: 10.1023/A:1003087532137 (1997) pdf

Ward, B. B., K. J. Courtney and J. H. Langenheim. Inhibition of Nitrosomonas europaea by monoterpenes from coastal redwood (Sequois sempervirens) in whole cell studies. Journal of Chemical Ecology, 23: 2583-2598. DOI: 10.1023/B:JOEC.0000006668.48855.b7 (1997) pdf

Ward, B. B., M.A. Voytek, K.-P. Witzel. Population diversity of ammonium oxidizers investigated by specific PCR amplification. Microbial Ecology, 33: 87-96. DOI: 10.1007/s002489900011 (1997)  pdf

Ward, B. B. Nitrification and denitrification: Probing the nitrogen cycle in aquatic environments. Microbial Ecology, 32: 247-261. DOI: 10.1007/BF00183061 (1996) pdf

Ward, B. B. Nitrification and ammonification in aquatic systems. Life Supp. Biosph. Sci., 3: 25-29. (1996)

Ward, B. B. Functional and taxonomic probes for bacteria in the nitrogen cycle. Ed. I. Joint, NATO workshop on Molecular Ecology of Aquatic Microbes, pp. 73-86. DOI: 10.1007/978-3-642-79923-5_5 (1995)  pdf

Ward, B. B. Diversity in denitrifying bacteria: Limits of DNA RFLP analysis and probes for the functional gene, nitrite reductase. Archives of Microbiology, 163: 167-175. DOI: 10.1007/BF00305349 (1995)  pdf

Voytek, M.A. and B. B. Ward. Detection of ammonium-oxidizing bacteria of the beta-subdivision proteobacteria in aquatic samples using the polymerase chain reaction. Applied and Environmental Microbiology, 61: 1441-1450. PMCID: PMC167402 (1995)  pdf

Bronk, D.A., P.M. Glibert and B. B. Ward. Nitrogen uptake, dissolved organic nitrogen release and new production. Science, 265: 1843-1846. DOI: 10.1126/science.265.5180.1843 (1994)  pdf

Miller, L.G. , M.D. Coutlakis, R.S. Oremland and B. B. Ward. Selective inhibition of nitrification (ammonium oxidation) by methylfluoride and dimethyl ether. Applied and Environmental Microbiology, 59: 2457-2464. PMCID: PMC182306 (1993)  pdf

Ward, B. B., A.R. Cockcroft and K.A. Kilpatrick. Antibody and DNA probes for detection of nitrite reductase in seawater. Journal of General Microbiology, 139: 2285-2293. DOI: 10.1099/00221287-139-9-2285 (1993)  pdf

Ward, B. B. and A.R. Cockcroft. Immunofluorescence detection of the denitrifying bacterium, Pseudomonas perfectomarina, in seawater and intertidal sediment environments. Microbial Ecology, 25: 233-246. DOI: 10.1007/BF00171890 (1993)  pdf

Ward, B. B. and K.A. Kilpatrick. Methane oxidation associated with mid-depth methane maxima in the Southern California Bight.  Continental Shelf Research, 13: 1111-1122. DOI: 10.1016/0278-4343(93)90044-X (1993)  pdf

Kerkhof, L.J. and B. B. Ward. Comparison of nucleic acid hybridization and fluorometry for measurement of RNA/DNA relationship with growth rate in a marine bacterium. Applied and Environmental Microbiology, 59: 1303-1309. PMCID: PMC182081 (1993)  pdf

Hansell, D.A., B. B. Ward and P.M. Williams. Measurements of DOC and DON in the Southern California Bight using oxidation by high temperature combustion.  Deep-Sea Research, 40: 219-234. DOI: 10.1016/0967-0637(93)90001-J (1993)  pdf

DeLong, E.F. and B. B. Ward. Biological oceanography from a molecular perspective. Oceanus 35: 47-54. (1992)

Ward, B. B. The subsurface methane maximum in the Southern California Bight. Continental Shelf Research, 12: 735-752. DOI: 10.1016/0278-4343(92)90028-I (1992)  pdf

Ward, B. B.   Nitrogen cycle of the sea. Encyclopedia of Earth System Science, Academic Press,Inc., 3:295-206. (1992)

Ward, B. B. and K.A. Kilpatrick. Nitrogen transformations in the oxic layer of permanent anoxic basins: The Black Sea and the Cariaco Trench. E. Izdar and J.W. Murray (eds.): Black Sea Oceanography, Kluwer Academic Publishers, The Netherlands, pp. 111-124. DOI: 10.1007/978-94-011-2608-3_7 (1991)   pdf

Reeburgh, W.S., B. B. Ward, S.C. Whalen, K.A. Sandbeck, K.A. Kilpatrick and L.J. Kerkhof. Black Sea Methane Geochemistry. Deep Sea Research. DOI: 10.1016/S0198-0149(10)80030-5 (1991)  pdf

Ward, B. B. Immunology in Biological Oceanography and Marine Ecology. The Oceanography Magazine, 3: 30-35. DOI: 10.5670/oceanog.1990.18 (1991)   pdf

Ward, B. B. and K.A. Kilpatrick. Relationship between substrate concentration and oxidation of ammonium and methane in a stratified water column. Continental Shelf Research, 10: 1193-1208. DOI: 10.1016/0278-4343(90)90016-F (1990)   pdf

Lipschultz, F., S.C. Wofsy, B. B. Ward, L.A. Codispoti, G. Friederich and J.W. Elkins. Bacterial transformations of inorganic nitrogen in the oxygen deficient waters of the eastern tropical south Pacific Ocean. Deep-Sea Research, 37: 1513-1541. DOI: 10.1016/0198-0149(90)90060-9 (1990)  pdf

Ward, B. B. Kinetics of ammonia oxidation by a marine nitrifying bacterium: Methane as a substrate analogue. Microbial Ecology, 19: 211-226. DOI: 10.1007/BF02017166 (1990)   pdf

Ward, B. B., H.E. Glover and F. Lipschultz. Chemoautotrophic activity and nitrifi­cation in the oxygen minimum zone off Peru. Deep-Sea Res., 36: 1031-1051.  DOI:10.1016/0198-0149(89)90076-9 (1989) pdf

Ward, B. B., K.A. Kilpatrick, E. Renger and R.W. Eppley. Biological nitrogen cycling in the nitracline. Limnol. Oceanogr., 34: 493-513.  DOI: 10.4319/lo.1989.34.3.0493 (1989) pdf

Spinrad, R.W., H.E. Glover, B. B. Ward, L. A. Codispoti and G. Kullenberg. Suspended particle and bacterial maxima in Peruvian coastal waters during a cold water anomaly. Deep-Sea Res., 36: 715-733. DOI: 10.1016/0198-0149(89)90147-7 (1989) pdf

Ward, B. B., K.A. Kilpatrick, A.E. Wopat, E.C. Minnich and M.E. Lidstrom. Methane oxidation in Saanich Inlet during summer stratification.  Continental Shelf Res., 9: 65-75. DOI:10.1016/0278-4343(89)90083-6 (1989) pdf

Ward, B. B. and O.C. Zafiriou. Nitrification and nitric oxide in the oxygen minimum of the eastern tropical North Pacific. Deep-Sea Res., 35: 1127- 1142. DOI:10.1016/0198-0149(88)90005-2 (1988) pdf

Heyman, U., B. Heyman and B. B. Ward. Cell affinity chromatography for a marine nitrifying bacterium. IN: C.M. Yentsch, F.C. Mague and P.K. Moran (Eds.), Immunochemical Approaches to Estuarine, Coastal and Oceanographic Questions, Springer-Verlag, 100-116. DOI: 10.1007/978-1-4684-7642-2_6 (1988)  pdf

Ward, B. B. Nitrogen transformations in the Southern California Bight. Deep-Sea Res., 34: 785-805. DOI: 10.1016/0198-0149(87)90037-9 (1987) pdf

Ward, B. B. Kinetic studies on ammonia and methane oxidation by Nitrosococcus Oceanus. Arch. Microbiol., 147: 126-133. DOI: 10.1007/BF00415273 (1987) pdf

Ward, B. B., K. A. Kilpatrick, P. C. Novelli and M. I. Scranton. Methane oxidation and methane fluxes in the ocean surface layer and in deep anoxic waters. Nature, 327: 226-229. DOI: 10.1038/327226a0  (1987) pdf

Ward, B. B.  Nitrification in Marine Environments, pp. 157-184. IN: J.I. Prosser (Ed.), Nitrification. Special Publications of the Society for General Microbiology, Vol. 20, IRL Press, Oxford. ISBN: 185221001X (1986)

Codispoti, L. I., G. E. Friederich, T. T. Packard, H. E. Glover, R. T. Barker, J. W. Elkins, B. B. Ward, F. Lipschultz and N. Lostaunau. High nitrite levels off northern Peru: A signal of instability in the marine denitrification rate. Science,233: 1200-1202. (1986) pdf

Ward, B. B. and A. F. Carlucci. Marine ammonium- and nitrite-oxidizing bacteria: Serological diversity determined by immunofluorescence in culture and in the environment. Appl. Environ. Microbiol., 50: 194-201. (1985) pdf

Ward, B. B. (Editor). Aquatic Nitrogen Cycles. Special Edition of Marine Chemistry. DOI: 10.1016/0304-4203(85)90050-7 (1985) pdf

Ward, B. B.  Light and substrate concentration effects on marine ammonium assimilation and oxidation rates. Mar., Chem., 16: 301-316. DOI: 10.1016/0304-4203(85)90052-0 (1985)  pdf

Ward, B. B., M. C. Talbot and M. J. Perry. Contributions of phytoplankton and nitrifying bacteria to ammonium and nitrite dynamics in coastal water.  Cont. Shelf Res., 3: 383-398. DOI: 10.1016/0278-4343(84)90018-9  (1984) pdf

Ward, B. B. Photosynthesis and bacterial utilization of phytoplankton exudates: Results from pre- and post-incubation size fractionation. Oceanol. Acta, 7: 337-343. https://archimer.ifremer.fr/doc/00131/24216/ (1984)  pdf

Karl, D.M., G. A. Knauer, J. H. Martin and B. B. Ward. Bacterial chemolithotrophy in the ocean is associated with sinking particles. Nature, 309: 54-56. DOI: 10.1038/309054a0 (1984) pdf

Ward, B. B. Autotrophic activity of ammonium-oxidizing bacteria: Combined autoradiography and immunofluorescence for estimation of single cell activity in the primary nitrite maximum off the coast of Washington. Limnol. Oceanogr., 29: 402-410. DOI: 10.4319/lo.1984.29.2.0402 (1984)  pdf

Ward, B. B. Oceanic distribution of ammonium-oxidizing bacteria determined by immunofluorescent assay. J. Mar. Res., 40: 1155-1172. (1982) pdf

Ward, B. B., R. J. Olson and M. J. Perry. Microbial nitrification rates in the primary nitrite maximum off Southern California. Deep-Sea Res., 29: 247-255. DOI: 10.1016/0198-0149(82)90112-1 (1982) pdf

Ward, B. B. and M. J. Perry. Immunofluorescent assay for the marine ammonium-oxidizing bacterium Nitrosococcus oceanus.  Appl. Environ. Microbiol, 39: 913-918. PMCID: PMC291443 (1980) pdf